In the distant future, a large portion of humanity lives on Mars, in cities that resemble those once found on Earth. The cities are protected from the inhospitable Martian atmosphere by dome-like force-fields. When a massive sandstorm breaks through the dome and destroys Mars New York, those in Mars Los Angeles must figure out how to stop the storm before it wipes them out next.

在萬(wàn)能兵器犯罪搜查官烏夫庫(kù)克博士（八島智人 配音）的協(xié)助下，潛入奢華無(wú)比但兇險(xiǎn)非常的賭場(chǎng)“樂(lè)園”的少女露恩·芭洛特（林原惠美 配音）經(jīng)過(guò)連番賭局，終于挫敗技術(shù)高潮的裝甲艾許雷，在籌碼上百萬(wàn)美金的賭局中獲勝，并且成功拿到藏有榭爾（中井和哉 配音）記憶的籌碼。可與此同時(shí)，鮑伊德（磯部勉 配音）緊追不舍，定要置露恩于死地。隨著榭爾的記憶大白天下，一切丑惡到了決斷的時(shí)刻……
本片根據(jù)沖方丁的科幻小說(shuō)改編，為“殼中少女”三部曲中最后一部（前兩部分別為“壓縮”和“燃燒”）。